Adule male of Pachydactylus atorquatus. © M. Barts
Few weeks old hatchling of Pachydactylus atorquatus. © M. Barts
Few weeks old hatchling of Pachydactylus atorquatus. © M. Barts
Habitat of Pachydactylus atorquatus. NW-South Africa. © M. Barts
Originalbeschreibung / Original description
BAUER, A., M. BARTS & F. HULBERT (2006): A new species of the Pachydactylus weberi group (Reptilia: Squamata: Gekkonidae) from the Orange River, with comments on its natural history. – Salamandra, Rheinbach, 42 (2/3): 83–92. — Terra typica: South Africa, Northern Cape Province, Augrabies National Park, Dassie Trail, in a sheltered depression on the plateau (28°35’15.6” S, 20°20’03.0” E, 619 m elevation).
Diagnosis: A large species, to 54.2 mm SVL (TM 84939), of the Pachydactylus weberi group. It may be distinguished from all other members of the P. weberi group by the com bination of the following characters: nasal region moderately inflated (strongly so in P. werneri); rostral, but not first supralabial nar rowly excluded from nostril (both excluded in P. kobosensis); dorsal scalation heteroge and a new species from the Karasburg District of Namibia); tail to at least 114 % of SVL, annulate, bearing whorls of slightly 4-6 enlarged, strongly keeled tubercles, each separated from others in same row by a single lanceolate scale (caudal tubercles abutting in P. fasciatus(dark, purplish-red in life) with no transverse body or nape bands (nape band only present in P. robertsi and a new species from the in all other members of the P. weberi group). Tips of tubercles on body and tail yellowish neous, with large, rounded to slightly oval, strongly keeled, juxtaposed tubercles ar ranged in 14 regular rows (partly imbricating in P. robertsi; tubercles in 20 or more rows in both P. robertsi and a new species from the Windhoek area); five undivided lamellae beneath digit IV of pes (typically six in P. acuminatus, P. kobosensis, P. tsodiloensis); adult dorsal pattern gray-brown Windhoek area; nape and body bands present to bright white (Figs. 2-5). A broad off-white stripe from nostril to orbit, bounded below by a dark stripe passing through loreal region and continuing behind the orbit to the level of the occiput; a bright white stripe beneath orbit. Juvenile pattern (Figs. 4-5) similar to adult (with transverse body and/or nape bands in all other species).
Description of holotype: Adult male. Snout-vent length (SVL) 54.2 mm. Body weakly depressed, moderately long (TrunkL/SVL ratio 0.43). Head relatively short (HeadL/SVL ratio 0.28), wide (HeadW/HeadL ratio 0.69), somewhat depressed (HeadH/HeadL ratio 0.41), clearly distinct from the neck. Lores and interorbital region weakly inflat ed. Snout long (Sn-Eye/HeadL ratio 0.39), longer than eye diameter (OrbD/Sn-Eye ratio 0.68); scales on snout and canthus enlarged relative to granular scales on crown, smooth, flattened to slightly domed; scales of interor bital and parietal regions strongly heteroge neous, with tiny granules interspersed with larger, domed to weakly conical, rounded to slightly oblong tubercles, each 30-100 % size of enlarged snout scales. Enlarged conical tu bercles regularly scattered across parietal and temporal regions, less prominent on occiput. Eye moderate (OrbD/HeadL ratio 0.26); or bits without extra-brillar fringes; approxi mately 7 supraciliary scales at posterodorsal corner of orbit bearing very small spines; pupil vertical, with crenellated margins. Ear opening oval, large (EarL/HeadL ratio 0.11); eye to ear distance greater than diameter of eyes (EyeEar/OrbD ratio 1.15). Rostral approximately 40% as deep (0.8 mm) as wide (2.0), no rostral groove, contacted by two enlarged supranasals and first suprala bials; nostrils oval, oriented dorsolaterally, each surrounded by two large postnasals, supranasal, and first supralabial; supranasals in contact anteriorly, separated by a single granule posteriorly; dorsal postnasals larger than ventral postnasals; nostril rims weakly inflated; one rows of scales separate orbit from supralabials; mental with nearly par allel sides, approximately 2.6 times deeper (2.3 mm) than wide (0.9 mm); no enlarged postmentals or chin shields. Enlarged su pralabials to angle of jaws 11/11 (8/8 to mid-orbit); infralabials 8/9; interorbital scale rows at midpoint of orbit 22 (12 across narrowest point of frontal bone).
Dorsal tubercles large (8-10 times size of adjacent scales), largest on midflanks and smallest along dorsal midline and across shoulders, rounded to slightly oblong, with a pronounced median keel – higher and broader posteriorly, forming 14 regular lon gitudinal rows on trunk, grading into coni cal granular scales on lower flanks; each tubercle surrounded by a rosette of small, irregular, conical scales of variable size; ven tral scales flattened, oval, subimbricate to imbricate, becoming somewhat larger pos teriorly, largest on posterior abdomen and in precloacal region, approximately 33 be tween lowest conical granular rows on flank at midbody; tubercular scales on dorsum at midbody much larger than ventral scales at same level; chin granules approximately one sixth to one fourth size of ventral scales, increasing in size behind the angle of jaws. No preanal or femoral pores. Scales on palm, sole, and ventral surface of forelimb smooth, flattened to granular; scales on ventral aspect of thighs enlarged, continuous with enlarged scales of precloacal region; scales on dorsal aspect of forelimb slightly domed to weakly conical, heterogeneous, subimbricate; scales on dorsum of thigh enlarged, conical, those on crus greatly enlarged.
Forelimbs short, stout (ForeaL/SVL ra tio 0.13); hindlimbs short (CrusL/SVL ratio 0.17); digits relatively long, claws not vis ible; subdigital scansors, except for distal most, entire, present only on distal portion of toes, approximately 1.5-2.0 times wider than more basal (non-scansorial) subdigital scales; interdigital webbing absent. Relative length of digits (manus): III>IV>V~II>I; (pes): IV>III~V>II>I. Subdigital scansors (excluding distal divided scansor) I (4), II (4), III (4), IV (4), V (4) – manus; I (4), II (5), III (5), IV (5), V (5) – pes.
Regenerated tail sub-cylindrical, short er than snout-vent length (TailL/SVL ratio 0.65); tail base with enlarged tubercles like those of body dorsum; regenerated portion with small, irregular, imbricating, smooth scales, much smaller on dorsum than on ventrum; some scales along ventral midline larger than adjacent scales; 2 small, smooth, dorsally-projecting postcloacal spurs on each side of hemipeneal bulge, anterior only slightly larger than posterior.
Coloration (in preservative): Dorsum a grayish brown, paler near limb insertions. Tubercles with tips or most of keel white. Dorsum of head with intergranular spaces whitish. A broad white stripe between nos tril and anterodorsal corner of orbit. Darker brown between white stripe and dorsal bor der of supralabials, on temporal region, and in circumauricular region, fading on nape. Infralabials and posterior supralabials cream colored with scattered brown pigment. Ven ter buff with scattered brown pigment around margins of flanks, throat, chin and limbs; palms, soles and digits mid-brown, midven tral region of chest and anterior abdomen pearly white; tail venter dusky.
Coloration (in life). The background col oration of the head, body and limbs is purple and the pupil is old rose (RAL 3014). The golden yellow (RAL 1004) and in the outerred (RAL 3004). A white (RAL 9016) stripe approximately three scale rows in width ex tends between the nostrils and the anterior margin of the eyes. A white (RAL 9016) stripe extends from the fourth supralabial, beneath the eye, to the posterior border of the ear. The loreal area between these two white stripes is reddish-black (RAL 3009) and a thick stripe of this color continues behind the orbit to the end of the lateral surface of the head. The supraoculars are darker than the dorsum of the head and may be character ized as brilliant blue (RAL 5007). The ciliary scales are zinc yellow (RAL 1018) dorsally and white (RAL 9016) around the remainder of the eye. The iris is violet blue (RAL 5000) tubercular scales of the dorsum and flanks are most rows the tips of the tubercles are a dis tinctly lighter pale gray (RAL 7035). On the dorsal surfaces of the forelimbs the tubercular scales are also golden yellow, whereas on the dorsal surfaces of the hindlimbs only the tips of the tubercles are this color. The tubercles of the tail are signal white (RAL 9003). The ventral and subcaudal scales are cream white (RAL 9001) and the areas between the scales are white (RAL 9016). The subdigital lamel lae are signal white (RAL 9003). Juveniles have essentially the same coloration as the adults, although the background coloration is somewhat lighter and the markings on the head are more distinct. A subadult male has a light blue (RAL 2012) sheen visible cau dally on the otherwise homogeneous typical coloration. Coloration varies with time of day in Pachydactylus atorquatus, with animals having a darker coloration by night.